Possible but I also suspect a php limit.
I’m going to test a download link without being logged in…
I confirm I can reproduce on a newly created instance. Works while authenticated, tiemouts when uploading through a shared link.
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I also can reproduce the issue now.
Not sure I understand the fix. But looking at the last posts of the GitHub issue I reported above they talk about an enhancement that will appear soon. It was a long lasting bug, 7 years 
The fix is for apache but we use nginx.
These could be the settings for nginx:
client_max_body_size 500m;
client_body_timeout 600;
Tried to edit ngnix.conf into the module :
# set max upload size (client_max_body_size as already set to 10G)
client_max_body_size 10G;
fastcgi_buffers 64 4K;
#MGT bigger timeout (for unauthenticated uploads)
client_body_timeout 600;
Not working 
If you look at the last posts of the GitHub issue I reported above they talk about an enhancement that will appear soon. It was a long lasting bug, 7 years
I suggest we wait for the fix.
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THANKS to @mrmarkus and others for helping nailing this nasty bug. I should have mentioned the quite unusual way I accessed the nextcloud drive (upload box link)
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