I decided to bypass some of the speculation and installed Wireshark on the DC and captured the LDAP traffic both when I perform and normal domain users and groups display on NS8 (successful) and when I retrieve User mailboxes on NS8 (fails). Here is a snippet of messages for the failing situation:
No. Time Source Destination Protocol Length Info
126 32.371949 192.168.1.38 192.168.1.2 LDAP 80 bindRequest(6) "<ROOT>" simple
No. Time Source Destination Protocol Length Info
127 32.372112 192.168.1.2 192.168.1.38 LDAP 88 bindResponse(6) success
No. Time Source Destination Protocol Length Info
129 32.372418 192.168.1.38 192.168.1.2 LDAP 286 searchRequest(5) "DC=DomainDnsZones,DC=burnsville,DC=local" wholeSubtree
No. Time Source Destination Protocol Length Info
130 32.372482 192.168.1.38 192.168.1.2 LDAP 286 searchRequest(3) "DC=ForestDnsZones,DC=burnsville,DC=local" wholeSubtree
No. Time Source Destination Protocol Length Info
131 32.372503 192.168.1.2 192.168.1.38 LDAP 240 searchResDone(5) operationsError (000004DC: LdapErr: DSID-0C09091F, comment: In order to perform this operation a successful bind must be completed on the connection., data 0, v2580) [0 results]
No. Time Source Destination Protocol Length Info
132 32.372605 192.168.1.2 192.168.1.38 LDAP 240 searchResDone(3) operationsError (000004DC: LdapErr: DSID-0C09091F, comment: In order to perform this operation a successful bind must be completed on the connection., data 0, v2580) [0 results]
I’m not sure where the missing bind is, but that seems to be why nothing is being returned.